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Wesener, T. (2016). The Giant Pill-Millipedes, order Sphaerotheriida – An annotated species catalogue with morphological atlas and list of apomorphies. Bonn zoological Bulletin Supplementum. 63: 1-107.
Wesener, T.
The Giant Pill-Millipedes, order Sphaerotheriida – An annotated species catalogue with morphological atlas and list of apomorphies
Bonn zoological Bulletin Supplementum
63: 1-107
CIM-ID: 7349
This is the first complete catalogue documenting all species, genera, and higher categories of the giant pill-millipedes, order Sphaerotheriida. It provides a list of all 39 known potential apomorphies of members of the order that could be used as a template for future phylogenetic studies at the ordinal level in the Diplopoda. The catalogue also includes illustrations of key characters of the order, as well as illustrations of the intraordinal variation of the female vulva and male posterior telopods. It further includes a key to all five Sphaerotheriida families, short descriptions for each of the 24 genera and a summary of the state of knowledge of the ecology and development of the giant pill-millipedes. Synonymies for all 24 valid genera are listed in a convenient tabular format. All 351 valid Sphaerotheriida species and their synonyms are listed; 14 remain nomina dubia, while five cannot be placed in any family. For each family, genus and species, bibliographic citations are presented, including all subsequent citations in chronological order. For each species, the number and location of type depositories as well as the locality data from the literature are provided. Bothrobelum sulcatulum (Butler, 1872) is presented as a new combination (ex Zephronia). Lectotypes are designated for Zoosphaerium blandum (deSaussure & Zehntner, 1902), Z. globulus & Z. imbecillum (both synonyms of Z. voeltzkowianum (deSaussure & Zehntner, 1897)), and Z. piligerum (deSaussure & Zehntner, 1897).
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2016-12-30 22:24:44Z
2023-06-26 10:26:03Z
2023-10-09 12:37:29Z

Arthrosphaeridae Jeekel, 1974 (additional source)
Borneopoeus Verhoeff, 1924 accepted as Castanotherium Pocock, 1895 (additional source)
Borneopoeus costatus Verhoeff, 1924 accepted as Castanotherium costatum (Verhoeff, 1924) (additional source)
Bothrobelum Verhoeff, 1924 (taxonomy source)
Bothrobelum rugosum Verhoeff, 1924 (taxonomy source)
Bothrobelum sulcatulum (Butler, 1872) (new combination reference)
Bournellum De Saussure & Zehntner, 1902 accepted as Sphaerotherium Brandt, 1833 (additional source)
Castanotherium amythrum (Attems, 1897) (additional source)
Castanotherium boetonense Carl, 1912 (additional source)
Castanotherium carinatum (Pocock, 1890) (additional source)
Castanotherium celebense Silvestri, 1897 (additional source)
Castanotherium cinctum (Carl, 1906) (taxonomy source)
Castanotherium conspicuum Silvestri, 1897 (additional source)
Castanotherium costatum (Verhoeff, 1924) (additional source)
Castanotherium crinitum (Attems, 1935) (additional source)
Castanotherium decoratum Carl, 1912 (additional source)
Castanotherium distinctum Carl, 1912 (additional source)
Castanotherium dorsispina (Attems, 1936) (additional source)
Castanotherium everettii Pocock, 1895 (additional source)
Castanotherium extraneum (Attems, 1943) (additional source)
Castanotherium falcicorne (Tömösvary, 1885) (taxonomy source)
Castanotherium ferum (Attems, 1935) (additional source)
Castanotherium glabratum (Newport, 1844) (additional source)
Castanotherium granulatum (Tömösvary, 1885) (taxonomy source)
Castanotherium hosei Pocock, 1895 (additional source)
Castanotherium humile (Silvestri, 1895) (additional source)
Castanotherium ignobile (Butler, 1872) (additional source)
Castanotherium indigenum (Attems, 1943) (additional source)
Castanotherium insigne (Brandt, 1833) (taxonomy source)
Castanotherium laeve Carl, 1912 (additional source)
Castanotherium lamprongse (Chamberlin, 1945) (taxonomy source)
Castanotherium moderatum (Chamberlin, 1945) (additional source)
Castanotherium nigriceps (Pocock, 1894) (additional source)
Castanotherium nigromaculatum Silvestri, 1897 (additional source)
Castanotherium obscurum Chamberlin, 1921 (additional source)
Castanotherium ornatum Carl, 1912 (additional source)
Castanotherium pellitum Attems, 1932 (additional source)
Castanotherium pilosum Carl, 1912 (additional source)
Castanotherium plasoni (Attems, 1943) (additional source)
Castanotherium porosum Pocock, 1895 accepted as Castanotheroides porosum (Pocock, 1895) accepted as Castanotheroides porosus (Pocock, 1895) (additional source)
Castanotherium pyrrhomelanum (Attems, 1897) (additional source)
Castanotherium rufipes (Pocock, 1894) (additional source)
Castanotherium simplex (Carl, 1909) (taxonomy source)
Castanotherium sparsepunctatum Carl, 1912 (additional source)
Castanotherium stellatum Carl, 1912 (additional source)
Castanotherium sulcicollis (Karsch, 1881) (additional source)
Castanotherium tivium (Chamberlin, 1945) (additional source)
Castanotherium tricollis (Karsch, 1881) (additional source)
Castanotherium variegatum (Attems, 1943) (additional source)
Castanotherium volzi (Carl, 1906) (additional source)
Castanotherium whiteheadi Pocock, 1895 (additional source)
Castanotheroides sinuatus (Butler, 1872) (additional source)
Chinosphaera Attems, 1936 accepted as Prionobelum Verhoeff, 1924 (additional source)
Chinosphaera maculosa Attems, 1935 accepted as Prionobelum maculosum (Attems, 1935) (additional source)
Cryxus ovalis (Linnaeus, 1758) (taxonomy source)
Cyliosoma Pocock, 1895 (additional source)
Cyliosoma angulatum (Butler, 1878) (additional source)
Cyliosomatidae Jeekel, 1974 (additional source)
Cyliosomella Verhoeff, 1924 accepted as Procyliosoma Silvestri, 1917 (additional source)
Epicyliosoma Silvestri, 1917 accepted as Cyliosoma Pocock, 1895 (additional source)
Eubournellum Attems, 1908 accepted as Sphaerotherium Brandt, 1833 (additional source)
Globotherium Brölemann, 1922 accepted as Zoosphaerium Pocock, 1895 (additional source)
Heligmasoma Chamberlin, 1921 accepted as Zoosphaerium Pocock, 1895 (additional source)
Indosphaera Attems, 1935 (additional source)
Indosphaera fontis Attems, 1936 (taxonomy source)
Kophosphaera Attems, 1935 (additional source)
Kophosphaera excavata (Butler, 1874) (taxonomy source)
Kophosphaera excavata mammifera Attems, 1936 accepted as Kophosphaera mammifera Attems, 1936 (additional source)
Kophosphaera mammifera Attems, 1936 (taxonomy source)
Kophosphaera pellita Attems, 1943 accepted as Sphaeropoeus pellitus (Attems, 1943) (taxonomy source)
Kophosphaera politissima Attems, 1936 (taxonomy source)
Leptoprotopus Silvestri, 1897 (additional source)
Leptoprotopus gladiator (Pocock, 1894) (taxonomy source)
Leptotelopus Silvestri, 1897 (taxonomy source)
Leptotelopus crepitans (Pocock, 1890) (taxonomy source)
Lissosphaera Attems, 1943 accepted as Sphaeropoeus Brandt, 1833 (additional source)
Lophozephronia Attems, 1936 accepted as Leptotelopus Silvestri, 1897 (additional source)
Luzonosphaera Wang, 1951 accepted as Castanotheroides Chamberlin, 1921 (additional source)
Microsphaerotherium Wesener & Vandenspiegel, 2007 (additional source)
Microsphaerotherium anjozorobe Wesener, 2009 (additional source)
Natalobelum Verhoeff, 1924 accepted as Sphaerotherium Brandt, 1833 (additional source)
Neobournellum Attems, 1908 accepted as Sphaerotherium Brandt, 1833 (additional source)
Odontosternum Attems, 1943 accepted as Arthrosphaera Pocock, 1895 (additional source)
Oligaspis Wood, 1865 accepted as Sphaerotherium Brandt, 1833 (additional source)
Pantitherium Attems, 1932 accepted as Sphaeropoeus Brandt, 1833 (additional source)
Paracyliosoma Verhoeff, 1928 accepted as Cyliosoma Pocock, 1895 (additional source)
Plethoconium Attems, 1943 accepted as Sphaerotheriida incertae sedis (additional source)
Prionobelum Verhoeff, 1924 (additional source)
Prionobelum durum Verhoeff, 1924 (additional source)
Procyliosoma Silvestri, 1917 (additional source)
Procyliosoma novazelandica (Kirk, 1886) (additional source)
Procyliosoma walesianum (Karsch, 1881) (taxonomy source)
Procyliosoma walesianum (Karsch, 1881) (basis of record)
Procyliosomatidae Wesener & VanDenSpiegel, 2009 (additional source)
Pulusphaera Attems, 1935 accepted as Castanotherium Pocock, 1895 (additional source)
Rajasphaera Attems, 1935 (additional source)
Rajasphaera montana Attems, 1935 (additional source)
Sphaerobelum Verhoeff, 1924 (taxonomy source)
Sphaerobelum clavigerum Verhoeff, 1924 (additional source)
Sphaerobelum truncatum Wongthamwanich, 2012 (taxonomy source)
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