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Hollier, J.; Stöckli, E.; Wesener, T.; Mesibov, R.; Decker, P.; Sierwald, P. (2020). An annotated list of the millipede (Diplopoda) species described by Johann Carl. Revue suisse de Zoologie. 127(1): 183-240.
381271
10.35929/RSZ.0015 [view]
Hollier, J.; Stöckli, E.; Wesener, T.; Mesibov, R.; Decker, P.; Sierwald, P.
2020
An annotated list of the millipede (Diplopoda) species described by Johann Carl
Revue suisse de Zoologie
127(1): 183-240
Publication
Johann Carl described 390 species or subspecies of millipedes during his long career at the Muséum d’histoire naturelle de Genève, studying specimens collected on his own expeditions and on those of other, mainly Swiss, naturalists. The type specimens deposited in Geneva are enumerated, and the type specimens that could be located in other collections in other museums are reported. Lectotypes are designated for 17 species: Brachyspirobolus aequatorialis (Carl, 1909), Carlopeltis alatus (Carl, 1914), Mastigorhacus alatus (Carl, 1912), Salpidobolus annulipes (Carl, 1912), Erythrhacus arietis (Carl, 1912), Argentocricus bernardinensis (Carl, 1918), Trigoniulus bitaeniatus Carl, 1912, Desmocricus conjunctus Carl, 1918, Polylepis elberti Carl, 1912, Trigoniulus incommodus Carl, 1912, Stenobolus insularis Carl, 1918, Salpidobolus lateralis (Carl, 1912), Dinematocricus lombokensis (Carl, 1912), Prepodesmus longipes (Carl, 1913), Arostrophus mertoni (Carl, 1912), Salpidobolus moenensis (Carl, 1912) and Eucarlia velox (Carl, 1912). Polydesmus carli nom. nov. is introduced as a replacement name for Polydesmus japonicus Carl, 1902, a junior homonym of Polydesmus japonicus Peters, 1864. Odontopyge johanncarli nom. nov. is introduced as a replacement name for Odontopyge fasciata Carl, 1905, a junior homonym of Odontopyge fasciata Attems, 1896.
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Agastrophus cinctellus Carl, 1926 accepted as Hypocambala cinctella (Carl, 1926) (additional source)
Agathodesmus baccatus (Carl, 1926) (additional source)
Amplinus convexus (Carl, 1902) (additional source)
Anoplodesmus cylindricus (Carl, 1935) (additional source)
Aphelidesmus ambiguus Carl, 1914 (additional source)
Argentocricus bernardinensis (Carl, 1918) (additional source)
Aulacobolus dysoni Carl, 1941 (additional source)
Aulacobolus ejaculans Carl, 1941 (additional source)
Aulacobolus ejaculans vallensis Carl, 1941 (additional source)
Batodesmus acceptus Carl, 1914 (additional source)
Brachyspirobolus aequatorialis (Carl, 1909) (additional source)
Brachyspirobolus domesticus (Carl, 1909) (additional source)
Calymmodesmus andinus Carl, 1914 (additional source)
Campodesmus costatus (Carl, 1913) (additional source)
Canacophilus acutangulus Carl, 1926 (additional source)
Carlopeltis alatus (Carl, 1914) (additional source)
Castanotherium boetonense Carl, 1912 (additional source)
Castanotherium cinctum (Carl, 1906) (additional source)
Castanotherium decoratum Carl, 1912 (additional source)
Castanotherium distinctum Carl, 1912 (additional source)
Chondrodesmus attemsi Carl, 1914 (additional source)
Chondrodesmus carbonarius Carl, 1914 (additional source)
Chondrodesmus convexus Carl, 1914 (additional source)
Chondrodesmus dorsovittatus Carl, 1914 (additional source)
Cingalobolus bugnioni Carl, 1918 (additional source)
Cryptocorypha areata (Carl, 1932) (additional source)
Cyrtodesmus ammonites (Carl, 1914) (additional source)
Desmocricus conjunctus Carl, 1918 (additional source)
Desmoxytes bicolor (Carl, 1902) (additional source)
Doratogonus annulipes Carl, 1917 (additional source)
Erythrhacus arietis (Carl, 1912) (additional source)
Geotypodon dispersus (Carl, 1909) (additional source)
Haplothysanus dorsalis (Carl, 1909) (additional source)
Harpagomorpha dentata (Carl, 1932) (additional source)
Harpurostreptus matarae Carl, 1941 (additional source)
Harpurostreptus montivagus Carl, 1941 (additional source)
Holopodostreptus braueri Carl, 1913 (additional source)
Hoplurorhachis crassipes (Carl, 1909) (additional source)
Hyleoglomeris alticola (Carl, 1912) (additional source)
Hypocambala caledonica (Carl, 1926) (additional source)
Ktenostreptus acceptus Carl, 1941 (additional source)
Lankabolus coelebs Carl, 1941 accepted as Lankabolus greeni (Pocock, 1892) (additional source)
Leptodesmus augustus Carl, 1914 (additional source)
Leptostreptus caudatus Carl, 1941 accepted as Leptostreptus caudiculatus (Karsch, 1881) (additional source)
Lophostreptus bicolor Carl, 1909 (additional source)
Lophostreptus ptilostreptoides Carl, 1909 (additional source)
Mastigorhacus alatus (Carl, 1912) (additional source)
Microspirobolus aequatorialis Carl, 1909 accepted as Bukobolus aequatorialis (Carl, 1909) accepted as Brachyspirobolus aequatorialis (Carl, 1909) (additional source)
Neocordyloporus asperus Carl, 1905 (additional source)
Nesorthomorpha armata (Carl, 1902) (additional source)
Nesorthomorpha coriacea (Carl, 1902) (additional source)
Nyssodesmus bivirgatus (Carl, 1902) accepted as Nyssodesmus phyton (Peters, 1864) (additional source)
Odontopyge johanncarli Hollier, Stöckli, Wesener, Mesibov, Decker & Sierwald, 2020 (status source)
Opisthoporodesmus bacillifer Carl, 1912 (additional source)
Orthomorpha butteli (Carl, 1922) (additional source)
Pagodesmus biporus Carl, 1932 (additional source)
Parabolus dimorphus (Carl, 1909) (additional source)
Parchondromorpha coonoorensis (Carl, 1932) (additional source)
Picrogonopus gracillimus Carl, 1941 (additional source)
Plethodesmus biseriatus Carl, 1926 (additional source)
Polydesmus americanus Carl, 1902 accepted as Pseudopolydesmus pinetorum (Bollman, 1888) (taxonomy source)
Polydesmus carli Hollier, Stöckli, Wesener, Mesibov, Decker & Sierwald, 2020 (status source)
Polydrepanum asperrimum (Carl, 1932) (additional source)
Polylepis elberti Carl, 1912 (additional source)
Poratophilus distinctus Carl, 1917 accepted as Zinophora distincta (Carl, 1917) (additional source)
Poratophilus junodi Carl, 1917 accepted as Zinophora junodi (Carl, 1917) (additional source)
Poratophilus similis Carl, 1917 accepted as Zinophora similis (Carl, 1917) (additional source)
Prepodesmus ornatus (Peters, 1864) (additional source)
Pseudopolydesmus pinetorum (Bollman, 1888) (additional source)
Pseudoprionopeltis cinereus Carl, 1902 (additional source)
Pseudosphaeroparia cardamomi Carl, 1932 (additional source)
Rhamphidarpe detruncata (Carl, 1905) (additional source)
Rhamphidarpe dorsosulcata (Carl, 1909) (additional source)
Rhinocricus carli Attems, 1914 accepted as Acladocricus carli (Attems, 1914) (additional source)
Rhinocricus colombianus Schubart, 1951 (additional source)
Rhinotus celebensis Carl, 1912 (additional source)
Rhinotus dempuranus Carl, 1912 (additional source)
Rhinotus ducalis Carl, 1926 (additional source)
Salpidobolus annulipes (Carl, 1912) (additional source)
Salpidobolus centralis (Carl, 1912) (additional source)
Salpidobolus centralis minor (Carl, 1912) (additional source)
Salpidobolus centralis spectabilis (Carl, 1912) (additional source)
Schizotelopus brevicornis (Carl, 1914) (additional source)
Sculptoteles braueri (Carl, 1918) (additional source)
Sholaphilus albidus Carl, 1932 (additional source)
Siphonophorella columbiana (Carl, 1914) (additional source)
Skotodesmus crepuscularis Carl, 1932 (additional source)
Solaenaulus butteli (Carl, 1922) (additional source)
Solaenaulus butteli birmanica Carl, 1941 (additional source)
Spirobolellus ambiguus Carl, 1926 (additional source)
Spirobolellus canalensis Carl, 1926 (additional source)
Spirobolellus comicus Carl, 1926 (additional source)
Spirobolellus dimidiatus Carl, 1926 (additional source)
Spirobolellus dispersus Carl, 1926 (additional source)
Spirobolellus duplus Jeekel, 2001 (additional source)
Spirostreptus angustifrons Carl, 1905 (additional source)
Spirostreptus frater Carl, 1909 accepted as Gonoplectus frater (Carl, 1909) (additional source)
Spirostreptus inconstans Carl, 1914 accepted as Hyloecostreptus inconstans (Carl, 1914) (additional source)
Spirostreptus malayus Carl, 1909 accepted as Gonoplectus malayus (Carl, 1909) (additional source)
Spirostreptus minor Carl, 1909 accepted as Thyropygus minor (Carl, 1909) accepted as Remulopygus minor (Carl, 1909) (additional source)
Unknown type BMNH, identified as Cryptocorypha areata (Carl, 1932)
Syntype SMF, identified as Trigoniulus bitaeniatus Carl, 1912
Syntype ZMHB, identified as Holopodostreptus braueri Carl, 1913
Syntype ZMHB, identified as Sculptoteles braueri (Carl, 1918)
Holotype ZMHB, verbatimGeounit Tjibodas, Zentraljav..., identified as Orthomorpha butteli (Carl, 1922)
Syntype ZMUH, identified as Odontopyge dorsalis Carl, 1909
Syntype ZMUH, identified as Microspirobolus domesticus Carl, 1909
Syntype MHNG MHNGARTO- 14250, identified as Steganostigmus contortipes Carl, 1932
Syntype MHNG MHNG-ARTO-14236, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14237, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14238, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14239, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14240, identified as Ktenostreptus acceptus Carl, 1941
Holotype MHNG MHNG-ARTO-14242, identified as Doratogonus annulipes Carl, 1917
Syntype MHNG MHNG-ARTO-14243, identified as Pagodesmus biporus Carl, 1932
Syntype MHNG MHNG-ARTO-14244, verbatimGeounit Palnis, Pumbarai and..., identified as Pagodesmus biporus Carl, 1932
Syntype MHNG MHNG-ARTO-14245, identified as Pagodesmus biporus Carl, 1932
Holotype MHNG MHNG-ARTO-14246, identified as Steganostigmus canonicus Carl, 1932
Holotype MHNG MHNG-ARTO-14247, identified as Pseudosphaeroparia cardamomi Carl, 1932
Paratype MHNG MHNG-ARTO-14248, identified as Pseudosphaeroparia cardamomi Carl, 1932
Syntype MHNG MHNG-ARTO-14249, identified as Steganostigmus contortipes Carl, 1932
Holotype MHNG MHNG-ARTO-14251, verbatimGeounit "Elizabethville (Kat..., identified as Poratophilus distinctus Carl, 1917
Holotype MHNG MHNG-ARTO-14252, identified as Xiphidiogonus dravidus Carl, 1932
Syntype MHNG MHNG-ARTO-14292, verbatimGeounit Neu-Caledonien: Mt. ..., identified as Canacophilus acutangulus Carl, 1926
Syntype MHNG MHNG-ARTO-14293, identified as Sundanina albicans (Carl, 1902)
Syntype MHNG MHNG-ARTO-14294, identified as Sholaphilus albidus Carl, 1932
Holotype MHNG MHNG-ARTO-14295, verbatimGeounit Soekaboemi, West-Jav..., identified as Tectoporus ambiguus (Carl, 1941)
Syntype MHNG MHNG-ARTO-14296, identified as Polydesmus americanus Carl, 1902
Syntype MHNG MHNG-ARTO-14312, identified as Salpidobolus centralis (Carl, 1912)
Lectotype MHNG MHNG-ARTO-14316, identified as Salpidobolus annulipes (Carl, 1912)
Syntype MHNG MHNG-ARTO-14322, verbatimGeounit Tchalabel N. Caled. ..., identified as Plethodesmus biseriatus Carl, 1926
Syntype MHNG MHNG-ARTO-14323, identified as Spirobolellus dimidiatus Carl, 1926
Syntype MHNG MHNG-ARTO-14327, verbatimGeounit Mt Canala Nlle. Cale..., identified as Spirobolellus canalensis Carl, 1926
Syntype MHNG MHNG-ARTO-14328, identified as Spirobolellus ambiguus Carl, 1926
Nontype MHNG MHNG-ARTO-14329, identified as Spirobolellus ambiguus Carl, 1926
Syntype MHNG MHNG-ARTO-14330, identified as Spirobolellus dispersus Carl, 1926
Syntype MHNG MHNG-ARTO-14331, verbatimGeounit N. Caled., Naltho, L..., identified as Spirobolellus dispersus Carl, 1926
Syntype MHNG MHNG-ARTO-14337, identified as Spirobolellus comicus Carl, 1926
Syntype MHNG MHNG-ARTO-14343, identified as Rhinotus ducalis Carl, 1926
Holotype MHNG MHNG-ARTO-14350, identified as Batodesmus acceptus Carl, 1914
Lectotype MHNG MHNG-ARTO-14351, verbatimGeounit Bodega Central (Colo..., identified as Carlopeltis alatus (Carl, 1914)
Syntype MHNG MHNG-ARTO-14353, identified as Calymmodesmus andinus Carl, 1914
Syntype MHNG MHNG-ARTO-14354, identified as Calymmodesmus andinus Carl, 1914
Syntype MHNG MHNG-ARTO-14355, identified as Trigoniulus andropygus impunctatus Carl, 1918
Syntype MHNG MHNG-ARTO-14356, identified as Trigoniulus andropygus impunctatus Carl, 1918
Syntype MHNG MHNG-ARTO-14357, identified as Trichomorpha annulipes Carl, 1914
Syntype MHNG MHNG-ARTO-14358, identified as Trichomorpha annulipes Carl, 1914
Syntype MHNG MHNG-ARTO-14359, identified as Leptodesmus augustus Carl, 1914
Holotype MHNG MHNG-ARTO-14360, identified as Nesorthomorpha armata (Carl, 1902)
Unknown type MHNG MHNG-ARTO-14362, verbatimGeounit Palnis inf., Tandiku..., identified as Polydrepanum asperrimum (Carl, 1932)
Lectotype MHNG MHNG-ARTO-14363, verbatimGeounit Maryland, Palnis inf..., identified as Polydrepanum asperrimum (Carl, 1932)
Syntype MHNG MHNG-ARTO-14364, verbatimGeounit Travancore Vattavada..., identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14365, verbatimGeounit Palnis sup. Vandarav..., identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14366, verbatimGeounit Palnis sup. Kukkal-S..., identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14367, identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14368, verbatimGeounit Palnis sup. Maryian-..., identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14369, verbatimGeounit Palnis sup. Kukkal, ..., identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14370, verbatimGeounit Palnis infér. Tandik..., identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14371, verbatimGeounit Palnis sup. Kodaikan..., identified as Cryptocorypha areata (Carl, 1932)
Lectotype MHNG MHNG-ARTO-14376, verbatimGeounit San Bernardino (Para..., identified as Argentocricus bernardinensis (Carl, 1918)
Lectotype MHNG MHNG-ARTO-14377, identified as Lophostreptus bicolor Carl, 1909
Paralectotype MHNG MHNG-ARTO-14378, identified as Lophostreptus bicolor Carl, 1909
Paralectotype MHNG MHNG-ARTO-14379, identified as Lophostreptus bicolor Carl, 1909
Lectotype MHNG MHNG-ARTO-14380, identified as Diglossosternum bipulvillatum (Carl, 1902)
Paralectotype MHNG MHNG-ARTO-14381, identified as Diglossosternum bipulvillatum (Carl, 1902)
Paralectotype MHNG MHNG-ARTO-14382, identified as Diglossosternum bipulvillatum (Carl, 1902)
Syntype MHNG MHNG-ARTO-14383, verbatimGeounit Costa Rica, St José,..., identified as Nyssodesmus bivirgatus (Carl, 1902)
Lectotype MHNG MHNG-ARTO-14388, identified as Parchondromorpha coonoorensis (Carl, 1932)
Paralectotype MHNG MHNG-ARTO-14389, identified as Parchondromorpha coonoorensis (Carl, 1932)
Lectotype MHNG MHNG-ARTO-14390, identified as Harpagomorpha dentata (Carl, 1932)
Paralectotype MHNG MHNG-ARTO-14391, identified as Harpagomorpha dentata (Carl, 1932)
Lectotype MHNG MHNG-ARTO-14401, identified as Orthomorpha coriacea Carl, 1902
Paralectotype MHNG MHNG-ARTO-14402, identified as Orthomorpha coriacea Carl, 1902
Syntype MHNG MHNG-ARTO-14403, identified as Polylepiscus braueri Carl, 1918
Syntype MHNG MHNG-ARTO-14404, verbatimGeounit Colombie, Bogota, 26..., identified as Stemmatoiulus bogotensis Carl, 1914
Syntype MHNG MHNG-ARTO-14405, verbatimGeounit Colombie, Bogota, 26..., identified as Stemmatoiulus bogotensis Carl, 1914
Syntype MHNG MHNG-ARTO-14406, verbatimGeounit Colombie, Bogota, 26..., identified as Stemmatoiulus bogotensis Carl, 1914
Syntype MHNG MHNG-ARTO-14407, verbatimGeounit Colombie, Bogota, 26..., identified as Stemmatoiulus bogotensis Carl, 1914
Syntype MHNG MHNG-ARTO-14408, verbatimGeounit Colombie, Bogota, 26..., identified as Stemmatoiulus bogotensis Carl, 1914
Syntype MHNG MHNG-ARTO-14409, verbatimGeounit Colombie, Bogota, 26..., identified as Stemmatoiulus bogotensis Carl, 1914
Holotype MHNG MHNG-ARTO-14410, identified as Cingalobolus bugnioni Carl, 1918
Holotype MHNG MHNG-ARTO-14411, identified as Solaenaulus butteli (Carl, 1922)
Holotype MHNG MHNG-ARTO-14413 and MHNG-ARTO-14414, identified as Cambalopsis butteli Carl, 1922
Lectotype MHNG MHNG-ARTO-14417, verbatimGeounit Njarugenje (Central-..., identified as Brachyspirobolus aequatorialis (Carl, 1909)
Unknown type MHNG MHNG-ARTO-14418, identified as Brachyspirobolus aequatorialis (Carl, 1909)
Syntype MHNG MHNG-ARTO-14419, identified as Microspirobolus domesticus Carl, 1909
Syntype MHNG MHNG-ARTO-14424, identified as Aulacobolus dysoni Carl, 1941
Syntype MHNG MHNG-ARTO-14425, identified as Aulacobolus dysoni Carl, 1941
Syntype MHNG MHNG-ARTO-14426, identified as Aulacobolus dysoni Carl, 1941
Syntype MHNG MHNG-ARTO-14427, verbatimGeounit Shola am Ibex-Hill b..., identified as Aulacobolus ejaculans Carl, 1941
Syntype MHNG MHNG-ARTO-14428, verbatimGeounit Hügel ob Nadu-Ar-Est..., identified as Aulacobolus ejaculans Carl, 1941
Syntype MHNG MHNG-ARTO-14429, identified as Aulacobolus ejaculans Carl, 1941
Syntype MHNG MHNG-ARTO-14430, verbatimGeounit Grosser Wald im ober..., identified as Aulacobolus ejaculans vallensis Carl, 1941
Syntype MHNG MHNG-ARTO-14431, identified as Aulacobolus ejaculans vallensis Carl, 1941
Unknown type MHNG MHNG-ARTO-14436, identified as Lankabolus coelebs Carl, 1941
Syntype MHNG MHNG-ARTO-14439, identified as Trigoniulus dimorphus Carl, 1909
Syntype MHNG MHNG-ARTO-14455, identified as Diopsiulus crassipes Carl, 1941
Syntype MHNG MHNG-ARTO-14456, identified as Diopsiulus crassipes Carl, 1941
Syntype MHNG MHNG-ARTO-14457, identified as Diopsiulus crassipes Carl, 1941
Syntype MHNG MHNG-ARTO-14458, identified as Diopsiulus crassipes Carl, 1941
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